[1] 张保海,姚学萍,王印,等.金黄色葡萄球菌Ⅶ型分泌系统研究进展[J].中国人兽共患病学报,2019,35(12): 1134-1138. [2] HOSPENTHAL M K,COSTA T R D,WAKSMAN G.A comprehensive guide to Pilus biogenesis in Gram-negative bacteria[J].Nat Rev Microbiol,2017,15(6): 365-379. [3] CARTER M E,BOGDANOVE A J,INNES R W,et al.A confounding effect of bacterial titer in a type III delivery-based assay of eukaryotic effector function[J].Mol Plant Microbe Interact,2018,31(11): 1115-1116. [4] MUÑOZ-GÓMEZ SA, HESS S, BURGER G, et al. An updated phylogeny of the Alphaproteobacteria reveals that the parasitic and have independent origins[J].Elife, 2019, 8: e42535. [5] DUPUIS L,MOREAU A,ROBERT D,et al.Structure and assembly of a bacterial nano-speargun: The type VI secretion system[J].Med Sci (Paris),2020,36(4): 408-412. [6] 谢立苹. 幽门螺杆菌CagM、CagI蛋白的生物信息学分析[D].镇江: 江苏大学,2012. [7] FRANCETIC O,BELIN D,BADAUT C,et al.Expression of the endogenous type II secretion pathway in Escherichia coli leads to chitinase secretion[J].EMBO J,2000,19(24): 6697-6703. [8] FERRANDEZ Y,CONDEMINE G.Novel mechanism of outer membrane targeting of proteins in Gram-negative bacteria[J].Mol Microbiol,2008,69(6): 1349-1357. [9] GAYTÁN M O,MARTÍNEZ-SANTOS V I,SOTO E,et al.Type three secretion system in attaching and effacing pathogens[J].Front Cell Infect Microbiol,2016,6: 129. [10] MILETIC S,GOESSWEINER-MOHR N,MARLOVITS T C.The structure of the type III secretion system needle complex[J].Curr Top Microbiol Immunol,2020,427: 67-90. [11] ZEPEDA-RIVERA M A,SAAK C C,GIBBS K A.A proposed chaperone of the bacterial type VI secretion system functions to constrain a self-identity protein[J].J Bacteriol,2018,200(14): e00688-e00617. [12] REDDY B L,SAIER M H JR.Properties and phylogeny of 76 families of bacterial and eukaryotic organellar outer membrane pore-forming proteins[J].PLoS One,2016,11(4): e0152733. [13] CHUNG JM, SHEEDLO MJ, CAMPBELL AM, et al.Structure of the Helicobacter pylori Cag type IV secretion system[J].Elife, 2019,8 :e47644. [14] JUHAS M,CROOK D W,HOOD D W.Type IV secretion systems: tools of bacterial horizontal gene transfer and virulence[J].Cell Microbiol,2008,10(12): 2377-2386. [15] LENTO C,FERRARO M,WILSON D,et al.HDX-MS and deletion analysis of the type 4 secretion system protein TraF from the Escherichia coli F plasmid[J].FEBS Lett,2016,590(3): 376-386. [16] SGRO G G,OKA G U,SOUZA D P,et al.Bacteria-killing type IV secretion systems[J].Front Microbiol,2019,10: 1078. [17] LI Y G,HU B,CHRISTIE P J.Biological and structural diversity of type IV secretion systems[M]//Protein Secretion in Bacteria.Washington,DC,USA: ASM Press,2019: 277-289. [18] Basler M,Pilhofer M,Henderson P G,et al.Type VI secretion requires a dynamic contractile phage tail-like structure[J]. Nature,2012,483(7388):182-186. [19] HAPFELMEIER S,DOMKE N,ZAMBRYSKI P C,et al.VirB6 is required for stabilization of VirB5 and VirB3 and formation of VirB7 homodimers in Agrobacterium tumefaciens[J].J Bacteriol,2000,182(16): 4505-4511. [20] MORSE K,NORIMINE J,PALMER G H,et al.Association and evidence for linked recognition of type IV secretion system proteins VirB9-1,VirB9-2,and VirB10 in Anaplasma marginale[J].Infect Immun,2012,80(1): 215-227. [21] GROHMANN E,CHRISTIE P J,WAKSMAN G,et al.Type IV secretion in Gram-negative and Gram-positive bacteria[J].Mol Microbiol,2018,107(4): 455-471. [22] GRAY T A,CLARK R R,BOUCHER N,et al.Intercellular communication and conjugation are mediated by ESX secretion systems in Mycobacteria[J].Science,2016,354(6310): 347-350. [23] GROHMANN E,KELLER W,MUTH G.Mechanisms of conjugative transfer and type IV secretion-mediated effector transport in gram-positive bacteria[J].Curr Top Microbiol Immunol,2017,413: 115-141. [24] 王俊平,郑玉玲,骈亚亚,等.猪链球菌2型Ⅳ型分泌系统组分VirD4与毒力相关性分析[J].微生物学报,2015,55(5): 643-649. [25] JUHAS M.Type IV secretion systems and genomic Islands-mediated horizontal gene transfer in Pseudomonas and Haemophilus[J].Microbiol Res,2015,170: 10-17. [26] TEGTMEYER N,WESSLER S,BACKERT S.Role of the cag-pathogenicity island encoded type IV secretion system in Helicobacter pylori pathogenesis[J].FEBS J,2011,278(8): 1190-1202. [27] AAS F E,WOLFGANG M,FRYE S,et al.Competence for natural transformation in Neisseria gonorrhoeae: components of DNA binding and uptake linked to type IV Pilus expression[J].Mol Microbiol,2002,46(3): 749-760. [28] 吴同垒,吴清民. 布鲁氏菌Ⅳ型分泌系统效应蛋白的研究进展[A]. 中国畜牧兽医学会家畜传染病学分会、解放军军事医学科学院军事兽医研究所.中国畜牧兽医学会家畜传染病学分会第八届全国会员代表大会暨第十五次学术研讨会论文集[C].中国畜牧兽医学会家畜传染病学分会、解放军军事医学科学院军事兽医研究所:中国畜牧兽医学会,2013:3. [29] KE Y H,WANG Y F,LI W F,et al.Type IV secretion system of Brucella spp.and its effectors[J].Front Cell Infect Microbiol,2015,5: 72. [30] CHANG Y W,SHAFFER C L,RETTBERG L A,et al.In vivo structures of the Helicobacter pylori cag type IV secretion system[J].Cell Rep,2018,23(3): 673-681. [31] STINGL K,MÜLLER S,SCHEIDGEN-KLEYBOLDT G,et al.Composite system mediates two-step DNA uptake into Helicobacter pylori[J].Proc Natl Acad Sci USA,2010,107(3): 1184-1189. [32] HARRISON O B,CLEMENCE M,DILLARD J P,et al.Genomic analyses of Neisseria gonorrhoeae reveal an association of the gonococcal genetic island with antimicrobial resistance[J].J Infect,2016,73(6): 578-587. [33] NAKANO N,KUBORI T,KINOSHITA M,et al.Crystal structure of Legionella DotD: insights into the relationship between type IVB and type II/III secretion systems[J].PLoS Pathog,2010,6(10): e1001129. [34] 杨长亮. 鲍曼不动杆菌生物膜形成的调节[J].中国感染控制杂志,2012,11(3): 228-230 [35] VAYSSIER-TAUSSAT M,LE RHUN D,DENG H K,et al.The Trw type IV secretion system of Bartonella mediates host-specific adhesion to erythrocytes[J].PLoS Pathog,2010,6(6): e1000946. |